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Arches National Park
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Ecology - Arches National Park
DESERTS
Arches lies near the heart of a desert called the “Colorado Plateau.” Deserts form when weather patterns or geographic land forms create an environment where lack of water limits biotic productivity. Water may exist in an unusable form such as ice, or may be absent altogether. There are four basic types of desert: high pressure, rain shadow, interior continental and coastal. High pressure deserts generally form at the middle latitudes (30 degrees) in each hemisphere where warm, dry air masses descend toward the earth's surface. Rain shadow deserts form in localized high pressure zones caused by warm, dry air descending from mountain ranges. The Colorado Plateau is also in the interior of a large continent, far away from significant water sources.
Because of the elevations throughout the region, with a mean of around 3,000 feet and peaks over 12,000 feet above sea level, the Colorado Plateau is also known as a cold or high desert. Though low humidity allows greater penetration of solar radiation, winter air temperatures frequently drop below freezing. In turn, summertime air and especially ground temperatures can reach levels lethal for many organisms. After sunset, the ground rapidly loses heat to the night sky and ambient air temperatures may drop significantly before dawn. Temperature fluctuations of over 40 degrees in a 24-hour period are not uncommon.
Arches receives more precipitation than many other deserts: about 9 inches annually. August is generally the wettest month, as weather systems from the southwest bring brief, intense tropical storms. However, precipitation is highly variable both temporally and spatially. During a single storm, one area may receive significantly more or less water than a neighboring spot less than a mile away.
SOIL COMPOSITION
Biological soil crust is a living groundcover that forms the foundation of high desert plant life in Arches and the surrounding area. This knobby, black crust is dominated by cyanobacteria, but also includes lichens, mosses, green algae, microfungi and bacteria.
Cyanobacteria, previously called blue-green algae, are one of the oldest known life forms. It is thought that these organisms were among the first land colonizers of the earth's early land masses, and played an integral role in the formation and stabilization of the earth's early soils. Extremely thick mats of these organisms converted the earth's original carbon dioxide-rich atmosphere into one rich in oxygen and capable of sustaining life.
When wet, Cyanobacteria move through the soil and bind rock or soil particles, forming an intricate web of fibers. In this way, loose soil particles are joined together, and an otherwise unstable surface becomes very resistant to both wind and water erosion. The soil-binding action is not dependent on the presence of living filaments. Layers of abandoned sheaths, built up over long periods of time, can still be found clinging tenaciously to soil particles, providing cohesion and stability in sandy soils at depths up to 10cm.
Nitrogen fixation is another significant capability of cyanobacteria. Vascular plants are unable to utilize nitrogen as it occurs in the atmosphere. Cyanobacteria are able to convert atmospheric nitrogen to a form plants can use. This is especially important in desert ecosystems, where nitrogen levels are low and often limiting to plant productivity.
Soil crusts have other functions as well, including an ability to intercept and store water, nutrients and organic matter that might otherwise be unavailable to plants.
EPHEMERAL POOLS
Throughout Arches, naturally occurring sandstone basins called "ephemeral pools" or "potholes" collect rain water and wind-blown sediment, forming tiny ecosystems where a fascinating collection of plants and animals have adapted to life in the desert. Potholes range from a few millimeters to a few meters in depth, and even the smallest potholes may harbor microscopic invertebrates.
To survive in a pothole, organisms must endure extreme fluctuations in several environmental factors. Surface temperatures vary from 140 degrees Fahrenheit in summer to below freezing in winter. As water evaporates, organisms must disperse to larger pools or tolerate dehydration and the drastic physical and chemical changes that accompany it.
The most extreme conditions exist when a pothole is dry. In addition to the wide temperature fluctuations, ultraviolet light from the sun can damage body tissues. Many aquatic organisms are adapted to acquiring oxygen through water and suffer when exposed to air. Pothole organisms have three main ways of dealing with drought.
"Drought escapers" are winged insects, amphibians and invertebrates that breed in potholes but cannot tolerate dehydration (e.g. mosquitoes, adult tadpole and fairy shrimp, spadefoot toads). In some cases, adults live in permanent water sources or on land and travel to temporary pools to mate and lay eggs. If the pool dries out before the young mature, they die. In the case of tadpole, fairy and clam shrimp, adults must lay their drought-tolerant eggs before the pool dries up.
"Drought resistors" (e.g. snails, mites) have a dormant stage resistant to drying out. These animals have a waterproof layer like a shell or exoskeleton that prevents body tissues from losing too much water while a pool is dry. By burrowing, these animals are able to seal themselves in the layers of fine mud that often coat the bottom of potholes and form an impermeable crust.
"Drought tolerators" (e.g. rotifers, tadpole and fairy shrimp eggs) are able to tolerate a loss of up to 92 percent of their total body water. This remarkable process, known as "cryptobiosis," is made even more remarkable by the fact that many cryptobiotic species can be rehydrated and become fully functional in as little as half an hour. Cryptobiosis is accomplished by a command center that remains hydrated while substituting sugar molecules for water throughout the rest of the body. This transfer maintains the structure and elasticity of an organism's cells during long periods of drought, and enables the organism to withstand the climatic extremes of the desert. In fact, brine shrimp have been hatched from cryptobiotic cysts that endured a flight on the outside of a spacecraft. Many tolerators have only one stage in their life cycle (e.g. egg, larva) that can survive desiccation, and will die if a pool dries up during another phase.
Pothole organisms not only have to endure dry spells, but also must evaluate conditions and decide when to break dormancy. Desert precipitation falls at irregular intervals, and once water enters a pothole there is no guarantee that there is enough for an organism to complete its life cycle. Most organisms living in potholes have very short life cycles, as brief as ten days, reducing the time water is required and allowing them to live in the shallow pools. Even vertebrates such as toads, which are found in other environments, display shorter development times when found in potholes.
However, the presence of water may not be the only cue used by eggs and dormant life forms to activate. Oxygen content, temperature, and other physical and chemical factors of the water may be evaluated. Some organisms produce different types of eggs that hatch on different cues; others lay eggs in different areas so that they experience slightly different environmental conditions. The net result is that not all eggs hatch at once and the species has a better chance of survival. After a pothole fills with water, the small ecosystem experiences many other changes. Water temperatures can be very high, while oxygen levels can be very low. As the pool shrinks from evaporation, its salinity increases and the pH changes. Many organisms are capable of surviving wide fluctuations in these factors, but for some these changes are an indication that the time for dormancy is near.
Arches lies near the heart of a desert called the “Colorado Plateau.” Deserts form when weather patterns or geographic land forms create an environment where lack of water limits biotic productivity. Water may exist in an unusable form such as ice, or may be absent altogether. There are four basic types of desert: high pressure, rain shadow, interior continental and coastal. High pressure deserts generally form at the middle latitudes (30 degrees) in each hemisphere where warm, dry air masses descend toward the earth's surface. Rain shadow deserts form in localized high pressure zones caused by warm, dry air descending from mountain ranges. The Colorado Plateau is also in the interior of a large continent, far away from significant water sources.
Because of the elevations throughout the region, with a mean of around 3,000 feet and peaks over 12,000 feet above sea level, the Colorado Plateau is also known as a cold or high desert. Though low humidity allows greater penetration of solar radiation, winter air temperatures frequently drop below freezing. In turn, summertime air and especially ground temperatures can reach levels lethal for many organisms. After sunset, the ground rapidly loses heat to the night sky and ambient air temperatures may drop significantly before dawn. Temperature fluctuations of over 40 degrees in a 24-hour period are not uncommon.
Arches receives more precipitation than many other deserts: about 9 inches annually. August is generally the wettest month, as weather systems from the southwest bring brief, intense tropical storms. However, precipitation is highly variable both temporally and spatially. During a single storm, one area may receive significantly more or less water than a neighboring spot less than a mile away.
SOIL COMPOSITION
Biological soil crust is a living groundcover that forms the foundation of high desert plant life in Arches and the surrounding area. This knobby, black crust is dominated by cyanobacteria, but also includes lichens, mosses, green algae, microfungi and bacteria.
Cyanobacteria, previously called blue-green algae, are one of the oldest known life forms. It is thought that these organisms were among the first land colonizers of the earth's early land masses, and played an integral role in the formation and stabilization of the earth's early soils. Extremely thick mats of these organisms converted the earth's original carbon dioxide-rich atmosphere into one rich in oxygen and capable of sustaining life.
When wet, Cyanobacteria move through the soil and bind rock or soil particles, forming an intricate web of fibers. In this way, loose soil particles are joined together, and an otherwise unstable surface becomes very resistant to both wind and water erosion. The soil-binding action is not dependent on the presence of living filaments. Layers of abandoned sheaths, built up over long periods of time, can still be found clinging tenaciously to soil particles, providing cohesion and stability in sandy soils at depths up to 10cm.
Nitrogen fixation is another significant capability of cyanobacteria. Vascular plants are unable to utilize nitrogen as it occurs in the atmosphere. Cyanobacteria are able to convert atmospheric nitrogen to a form plants can use. This is especially important in desert ecosystems, where nitrogen levels are low and often limiting to plant productivity.
Soil crusts have other functions as well, including an ability to intercept and store water, nutrients and organic matter that might otherwise be unavailable to plants.
EPHEMERAL POOLS
Throughout Arches, naturally occurring sandstone basins called "ephemeral pools" or "potholes" collect rain water and wind-blown sediment, forming tiny ecosystems where a fascinating collection of plants and animals have adapted to life in the desert. Potholes range from a few millimeters to a few meters in depth, and even the smallest potholes may harbor microscopic invertebrates.
To survive in a pothole, organisms must endure extreme fluctuations in several environmental factors. Surface temperatures vary from 140 degrees Fahrenheit in summer to below freezing in winter. As water evaporates, organisms must disperse to larger pools or tolerate dehydration and the drastic physical and chemical changes that accompany it.
The most extreme conditions exist when a pothole is dry. In addition to the wide temperature fluctuations, ultraviolet light from the sun can damage body tissues. Many aquatic organisms are adapted to acquiring oxygen through water and suffer when exposed to air. Pothole organisms have three main ways of dealing with drought.
"Drought escapers" are winged insects, amphibians and invertebrates that breed in potholes but cannot tolerate dehydration (e.g. mosquitoes, adult tadpole and fairy shrimp, spadefoot toads). In some cases, adults live in permanent water sources or on land and travel to temporary pools to mate and lay eggs. If the pool dries out before the young mature, they die. In the case of tadpole, fairy and clam shrimp, adults must lay their drought-tolerant eggs before the pool dries up.
"Drought resistors" (e.g. snails, mites) have a dormant stage resistant to drying out. These animals have a waterproof layer like a shell or exoskeleton that prevents body tissues from losing too much water while a pool is dry. By burrowing, these animals are able to seal themselves in the layers of fine mud that often coat the bottom of potholes and form an impermeable crust.
"Drought tolerators" (e.g. rotifers, tadpole and fairy shrimp eggs) are able to tolerate a loss of up to 92 percent of their total body water. This remarkable process, known as "cryptobiosis," is made even more remarkable by the fact that many cryptobiotic species can be rehydrated and become fully functional in as little as half an hour. Cryptobiosis is accomplished by a command center that remains hydrated while substituting sugar molecules for water throughout the rest of the body. This transfer maintains the structure and elasticity of an organism's cells during long periods of drought, and enables the organism to withstand the climatic extremes of the desert. In fact, brine shrimp have been hatched from cryptobiotic cysts that endured a flight on the outside of a spacecraft. Many tolerators have only one stage in their life cycle (e.g. egg, larva) that can survive desiccation, and will die if a pool dries up during another phase.
Pothole organisms not only have to endure dry spells, but also must evaluate conditions and decide when to break dormancy. Desert precipitation falls at irregular intervals, and once water enters a pothole there is no guarantee that there is enough for an organism to complete its life cycle. Most organisms living in potholes have very short life cycles, as brief as ten days, reducing the time water is required and allowing them to live in the shallow pools. Even vertebrates such as toads, which are found in other environments, display shorter development times when found in potholes.
However, the presence of water may not be the only cue used by eggs and dormant life forms to activate. Oxygen content, temperature, and other physical and chemical factors of the water may be evaluated. Some organisms produce different types of eggs that hatch on different cues; others lay eggs in different areas so that they experience slightly different environmental conditions. The net result is that not all eggs hatch at once and the species has a better chance of survival. After a pothole fills with water, the small ecosystem experiences many other changes. Water temperatures can be very high, while oxygen levels can be very low. As the pool shrinks from evaporation, its salinity increases and the pH changes. Many organisms are capable of surviving wide fluctuations in these factors, but for some these changes are an indication that the time for dormancy is near.
